Tits and Chickadees
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Great Tit
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Scientific classification
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Kingdom:
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Animalia
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Phylum:
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Chordata
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Class:
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Aves
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Order:
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Passeriformes
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Family:
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Paridae
Vigors, 1825
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Genera
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5-10, see text.
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Synonyms
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see text
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The tits, chickadees,
and titmice, family Paridae, are a large family of small passerine
birds which occur in the northern hemisphere and Africa. Most were formerly in
the genus Parus.
These birds are called
"chickadees" (onomatopoeic, derived from their distinctive
"chick-a dee dee dee" alarm call) or "titmice" in North
America, and just "tits" in the rest of the English speaking world. The
name titmouse is attested from the 14th century, composed of the Old
English name for the bird, mase (Proto-Germanic *maison) and tit,
denoting something small. The spelling was influenced by mouse in the 16th
century.
These birds are mainly
small stocky woodland species with short stout bills. Some have crests. They
are adaptable birds, with a mixed diet including seeds and insects. Many
species will live around human habitation and come readily to bird feeders for
nuts or seed, and learn to take other foods. In England, Great Tits and Blue
Tits famously learned to break open the foil caps sealing bottles of milk that
had been delivered to homes to get at the cream floating on top.
These are hole-nesting
birds laying speckled white eggs.
Systematics
More recently, the large Parus
group has been gradually split into several genera (as indicated below), which
has been pioneered by North American ornithological authorities and to a more
limited degree (as of now) elsewhere. Whereas in the mid-1990s, only Pseudopodoces,
Baeolophus, Melanochlora and Sylviparus were considered
well-supported by the available data as distinct from Parus (Harrap
& Quinn 1996). Today, this arrangement is considered paraphyletic as
indicated by mtDNA cytochrome b sequence analysis and Parus is
best restricted to the Parus major - Parus fasciiventer clade,
and even the latter species' closest relatives might be considered a distinct
genus (Gill et al. 2005).
In the Sibley-Ahlquist
taxonomy, the Paridae family is much enlarged to include related groups such as
the Penduline tits and Long-tailed tits, but while the former are quite close
to the titmice indeed and could conceivably included in that family together with
the stenostirid "warblers", the long-tailed tits are not. Indeed, the
Yellow-browed Tit and the Sultan Tit are possibly more distant to the titmice
than are the penduline tits (Gill et al. 2005 and Jønsson & Fjeldsa
2006). If the two current families are lumped into the Paridae, the titmice
would be a subfamily Parinae.
Alternatively, all tits -
save the 2 monotypic genera discussed in the preceding section and possibly Cyanistes,
but including Hume's Ground Tit - could be lumped in Parus. In any case,
4 major clades of "typical" tits can be recognized: the dark-capped
chickadees and their relatives (Poecile including Sittiparus),
the long-crested Baeolophus and Lophophanes species, the usually
tufted, white-cheeked Periparus (including Pardaliparus) with
more subdued coloration and finally Parus sensu stricto (including Melaniparus
and Macholophus). Still, the interrelationship of these as well as the
relationships of many species within the clades are not well resolved at all;
analysis of morphology and biogeography probably gives more a robust picture
than the available molecular data (Gill et al., 2005).
Titmice have settled North
America twice, probably at some time during the Early-Mid Pliocene. The first
were the ancestors of Baeolophus; chickadees arrived somewhat later
(Gill et al., 2005).
FAMILY PARIDAE
- Penduline tits and
- stenostirid "warblers"
might be included here
- Genus Sylviparus
- Yellow-browed Tit, Sylviparus modestus
- Genus Melanochlora
- Sultan Tit, Melanochlora sultanea
These two monotypic genera are possibly less
close to titmice than are the penduline tits.
- Genus Cyanistes - frequently included in Parus
- Blue Tit, Cyanistes
caeruleus
- Azure Tit, Cyanistes
cyanus
- Yellow-breasted Tit, Cyanistes
flavipectus
- Genus Baeolophus
- Bridled Titmouse, Baeolophus
wollweberi
- Oak Titmouse, Baeolophus inornatus
- Juniper Titmouse, Baeolophus ridgwayi
- Tufted Titmouse, Baeolophus bicolor
- Black-crested Titmouse, Baeolophus
atricristatus
- Genus Lophophanes - frequently included in Parus
- Crested Tit, Lophophanes cristatus
- Grey-crested Tit, Lophophanes dichrous
- Genus Periparus - frequently included in Parus
- Rufous-naped Tit or Black-breasted Tit, Periparus
rufonuchalis
- Rufous-vented Tit, Periparus
rubidiventris
- Palawan Tit, Periparus amabilis -
sometimes separated in Pardaliparus
- Yellow-bellied Tit, Periparus
venustulus - sometimes separated in Pardaliparus
- Elegant Tit, Periparus elegans -
probably paraphyletic; sometimes separated in Pardaliparus
- Coal Tit, Periparus ater -
possibly paraphyletic
- Spot-winged Tit or Black-crested Tit, Periparus
ater melanolophus
- Genus Poecile - frequently included in Parus
- White-browed Tit, Poecile superciliosa
- Sombre Tit, Poecile
lugubris
- Varied Tit, Poecile varia -
sometimes separated in Sittiparus
- Daito Varied Tit, Poecile varia orii
- extinct (c.1940s)
- White-fronted Tit, Poecile semilarvata
- sometimes separated in Sittiparus
- Caspian Tit, Poecile
hyrcana
- Père David's Tit, Poecile
davidi
- Marsh Tit, Poecile
palustris
- Black-bibbed Tit, Poecile (palustris)
hypermelaena
- Willow Tit, Poecile montana
- Songar Tit, Poecile
songara
- Carolina Chickadee, Poecile
carolinensis
- Mexican Chickadee, Poecile sclateri
- Black-capped Chickadee, Poecile
atricapillus
- Mountain Chickadee, Poecile gambeli
- Siberian Tit or Gray-headed Chickadee, Poecile
cincta
- Boreal Chickadee, Poecile hudsonica
- Chestnut-backed Chickadee, Poecile
rufescens
- Genus Parus
- Carp's Tit, Parus carpi -
sometimes separated in Melaniparus
- White-bellied Tit, Parus albiventris
- sometimes separated in Melaniparus
- White-shouldered Tit, Parus guineensis
- sometimes separated in Melaniparus
- White-winged Black Tit, Parus leucomelas
- sometimes separated in Melaniparus
- Southern Black Tit, Parus niger -
sometimes separated in Melaniparus
- White-backed Tit, Parus leuconotus
- sometimes separated in Melaniparus
- Dusky Tit, Parus funereus -
sometimes separated in Melaniparus
- Rufous-bellied Tit, Parus rufiventris
- sometimes separated in Melaniparus
- Cinnamon-breasted Tit, Parus
pallidiventris - sometimes separated in Melaniparus
- Red-throated Tit, Parus fringillinus
- sometimes separated in Melaniparus
- Stripe-breasted Tit, Parus
fasciiventer - sometimes separated in Melaniparus
- Acacia Tit or Somali Tit, Parus
thruppi - sometimes separated in Melaniparus
- Miombo Tit, Parus griseiventris -
sometimes separated in Melaniparus
- Ashy Tit, Parus cinerascens -
sometimes separated in Melaniparus
- Southern Grey Tit, Parus afer -
sometimes separated in Melaniparus
- Yellow Tit, Parus holsti -
sometimes separated in Macholophus
- Yellow-cheeked Tit, Parus spilonotus
- Black-lored Tit, Parus xanthogenys
- sometimes separated in Macholophus
- Great Tit, Parus major
- Japanese Tit, Parus
minor
- Turkestan Tit, Parus
bokharensis
- Green-backed Tit, Parus monticolus
- White-winged Tit, Parus nuchalis
- Genus Pseudopodoces
- Hume's Ground Tit, previously
"Hume's Ground Jay", Pseudopodoces humilis
This
species has only recently been removed from the Crow family Corvidae and placed
here[citation needed].
References
- Gill, Frank B.; Slikas, Beth & Sheldon,
Frederick H. (2005): Phylogeny of titmice (Paridae): II. Species
relationships based on sequences of the mitochondrial cytochrome-b gene. Auk
122: 121-143. DOI:
10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2 HTML abstract
- Harrap, Simon & Quinn, David (1996): Tits,
Nuthatches & Treecreepers. Christopher Helm, London. ISBN
0-7136-3964-4
- Jønsson, Knud A. & Fjeldså, Jon
(2006): Determining biogeographical patterns of dispersal and
diversification in oscine passerine birds in Australia, Southeast Asia and
Africa. J. Biogeogr. 33(7): 1155–1165. DOI:10.1111/j.1365-2699.2006.01507.x (HTML abstract)