Scientific
classification
Kingdom: |
Animalia |
Phylum: |
Chordata |
Class: |
Mammalia |
Order: |
Cetacea |
Suborder: |
Odontoceti |
Family: |
Delphinidae |
The name is from Ancient
Greek δελφίς delphis meaning "with a womb" which can be interpreted as meaning
"a 'fish' with a womb".[1]
The word is used in a few
different ways. It can mean:
In this article, the second
definition is used. Porpoises (suborder Odontoceti, family Phocoenidae) are
thus not dolphins in this sense. Orcas and some closely related species belong
to the Delphinidae family and therefore qualify as dolphins, even though they are
called whales in common language. A group of dolphins can be called a
"school" or a "pod".
Six species in the family
Delphinidae are commonly called "whales" but are strictly speaking
dolphins. They are sometimes called "blackfish".
In 1933, three abnormal
dolphins were beached off the Irish coast; these appeared to be hybrids between
Risso's Dolphin and the Bottlenose Dolphin.[2] This mating has since
been repeated in captivity and a hybrid calf was born. In captivity, a
Bottlenose Dolphin and a Rough-toothed Dolphin produced hybrid offspring.[3]
A Common-Bottlenose hybrid lives at SeaWorld California.[4] Various
other dolphin hybrids have also been reported in the wild, such as a
Bottlenose-Atlantic Spotted hybrid.[5]
Dolphins, along with whales
and porpoises, are thought to be descendants of terrestrial mammals, most
likely of the Artiodactyl order. Dolphins entered the water roughly fifty
million years ago. Dolphins have a streamlined fusiform body, adapted for fast
swimming. The basic coloration patterns are shades of gray with a light
underside and a distinct dark cape on the back. It is often combined with lines
and patches of different hue and contrast.
Modern dolphin skeletons
have two small, rod-shaped pelvic bones thought to be vestigial hind legs. In
October of 2006, an unusual Bottlenose Dolphin was captured in Japan that had
small fins on either side of the genital slit, which scientists believe to be a
more pronounced development of these vestigal hind legs.[6]
The head contains the
melon, a round organ used for echolocation. In many species, the jaws are
elongated, forming a distinct beak; for some species like the Bottlenose, there
is a curved mouth that looks like a fixed smile. Teeth can be very numerous (up
to two hundred and fifty) in several species.
The dolphin brain is large
and has a highly structured cortex, which often is referred to in discussions
about their advanced intelligence. A recent theory however disputes the
existence of a neural basis for dolphin intellect, suggesting instead that the
large brain is only an adaptation to living in cold water.[7]
Unlike most mammals,
dolphins do not have hair, but they are born with a few hairs around the tip of
their rostrum which they lose after some time, in some cases even before they
are born. River dolphins usually have them all their life and are believed to
have a sensory function. Even in those species that lose the hair the hair
follicles remain, and it is possible the nerves connected to them are still
able to detect movements in the water.
Most dolphins have acute
eyesight, both in and out of the water, and their sense of hearing is superior
to that of humans. Though they have a small ear opening on each side of their
head, it is believed that hearing underwater is also if not exclusively done
with the lower jaw which conducts the sound vibrations to the middle ear via a
fat-filled cavity in the lower jaw bone. Hearing is also used for echolocation,
which seems to be an ability all dolphins have. Their teeth are arranged in a
way that works as an array or antenna to receive the incoming sound and make it
easier for them to pinpoint the exact location of an object.[8] The
dolphin's sense of touch is also well-developed. However, dolphins lack an
olfactory nerve and lobes and thus are believed to have no sense of smell,[9]
but they can taste and do show preferences for certain kinds of fish. Since
dolphins spend most of their time below the surface normally, just tasting the
water could act in a manner analogous to a sense of smell.
The Anatomy
of a Dolphin showing its skeleton, major organs and body shape.
Hind
Limb Buds on Dolphins
An embryo of a Spotten Dolphin in the fifth week of development. The hind limbs
are present as small bumps (hind limb buds) near the base of the tail. The pin
is approximately 1 inch long. Image used with permission, courtesy Thewissen
LabSource [1]
Dolphins are often regarded
as one of Earth's most intelligent species, though it is hard to say just how
intelligent dolphins are as straightforward comparisons of species' relative
intelligence are complicated by differences in sensory apparatus, response
modes, and nature of cognition. Furthermore, the difficulty and expense of doing
experimental work with large aquatics means that some tests that could
meaningfully be done still have not been carried out, or have been carried out
with inadequate sample size and methodology. See the "cetacean
intelligence" article for more details.
Dolphins often leap above
the water surface, sometimes performing acrobatic figures (e.g. the spinner
dolphin). Scientists are not quite certain about the purpose of this behavior,
but it may be to locate schools of fish by looking at above-water signs, like
feeding birds. They could also be communicating to other dolphins to join a
hunt, attempting to dislodge parasites, or simply doing it for fun. Play is a
very important part of dolphins' lives, and they can often be observed playing
with seaweed or play-fighting with other dolphins. They even harass other
locals, like seabirds and turtles. Dolphins also seem to enjoy riding waves and
frequently 'surf' coastal swells and the bow waves of boats. Dolphins are also
known to engage in acts of aggression towards each other. The older a male
dolphin is, the more likely his body is covered with scars ranging in depth
from teeth marks made by other dolphins. It is suggested that male dolphins
engage in such acts of aggression for the same reasons as humans: disputes
between companions or even competition for other females. Acts of agression can
become so intense that targeted dolphins are known to go into exile: leave
their communities as a result of losing a fight with other dolphins.
They are also willing to
occasionally approach humans and playfully interact with them in the water. In
return, some human cultures such as the Ancient Greeks treated them with
welcome; a ship spotting dolphins riding in their wake was considered a good
omen for a smooth voyage. There are many stories of dolphins protecting
shipwrecked sailors against sharks by swimming circles around them.
Dolphins are social, living
in pods (also called "schools") of up to a dozen individuals. In
places with a high abundance of food, pods can join temporarily, forming an
aggregation called a superpod; such groupings may exceed a thousand
dolphins. The individuals communicate using a variety of clicks, whistles and
other vocalizations. They also use ultrasonic sounds for echolocation.
Membership in pods is not rigid; interchange is common. However, the cetaceans
can establish strong bonds between each other. This leads to them staying with
injured or ill individuals for support.
Because of their capacity
for learning, dolphins have been employed by humans for any number of purposes.
Dolphins trained to perform in front of an audience have become a favorite
attraction in dolphinaria, for example SeaWorld. Such places may sometimes also
provide an opportunity for humans to interact very closely with dolphins.
Dolphin-human interaction is also employed in a curative sense at places where
dolphins work with autistic or otherwise disabled human children. The military
has employed dolphins for various purposes from finding mines to rescuing lost
or trapped humans. Such military dolphins, however, drew scrutiny during the
Vietnam War when rumors circulated that dolphins were being trained to kill
Vietnamese skin divers. Reports of cooperative human-dolphin fisheries date
back to Pliny. A modern human-dolphin fishery was reported in Laguna, Santa
Catarina, Brazil in 1990.
In May 2005, researchers in
Australia discovered a cultural aspect of dolphin behaviour: Some dolphins (Tursiops
aduncus) teach their children to use tools. The dolphins break sponges off
and cover their snouts with them thus protecting their snouts while foraging.
This knowledge of how to use a tool is mostly transferred from mothers to
daughters, unlike simian primates, where the knowledge is generally passed onto
all the young, irrespective of sex. The technology to use sponges as mouth
protection is not genetically inherited but a taught behaviour.[10]
Dolphins are one of the few
animals other than humans known to mate for reasons other than reproduction and
especially male Bottlenose Dolphins are known to engage in sexual acts with
other dolphin species, which is not always consensual, though the Bottlenose
may also be submissive in such encounters.[11] Occasionally,
dolphins will also show sexual behaviour towards humans.[12]
Male dolphins have been
known to engage in infanticide. Dolphins will also kill porpoises for reasons
which are not fully understood, as porpoises generally do not share the same
fish diet as dolphins and are therefore not competitors for food supplies.[13]
Individual species may
employ a number of methods of hunting. One such method is herding, where a
superpod will control a school of fish while individual members take turns
plowing through the herd, feeding. The tightly packed school of fish is
commonly known as bait ball. Coralling is a method where fish are chased to
shallow water where they are more easily captured. In South Carolina, coastal
Bottlenose Dolphins take this one step further with what has become known as
mudding, where the fish are driven onto mud banks and retrieved from there. In
some places, Orcas will also come up to the beach to capture Seals. Some
species also whack fish with their fluke, stunning them and sometimes sending
fish clear out of the water.
Some dolphin species face
an uncertain future, including pink dolphins, black dolphins, Amazon River
dolphins, and the Ganges and Yangtze River dolphins, all of which are
critically or seriously endangered. A 2006 survey found no individuals of the
Yangtze River dolphin, leading to fears that the species may already be
extinct.[14]
Contamination of
environment - the oceans, seas, and rivers - is an issue of concern, especially
pesticides, heavy metals, plastics, and other industrial and agricultural
pollutants that do not disintegrate rapidly in the environment are reducing
dolphin populations, and resulting in dolphins building up unusually high
levels of contaminants. Injuries or deaths due to collisions with boats,
especially their propellers, are also not uncommon.
Various fishing methods,
most notably purse seine fishing for tuna and the use of drift and gill nets,
results in a large amounts of dolphins being killed inadvertently.[15]
Accidental by-catch in trout nets is also not uncommon and poses a risk for
mainly local dolphin populations. In some parts of the world, such as some
areas in Japan and the Faroe Islands, dolphins are traditionally considered as
food, and killed in harpoon or drive hunts.
The year 2007 has been
declared as (International) Year of the Dolphin by the United Nations and
United Nations Environment Programme (UNEP).[16]
The UN Convention on
Migratory Species, together with its specialized agreements on dolphin
conservation ACCOBAMS and ASCOBANS and the WDCS (Whale and Dolphin Conservation
Society) have proposed 2007 as the "Year of the Dolphin" ('YOD')).
Dolphins have long played a
role in human culture. Dolphins are common in Greek mythology and there are
many coins from the time that feature a man or boy riding on the back of a
dolphin. Dolphins also seem to have been important to the Minoans, judging by
artistic evidence from the ruined palace at Knossos. In Hindu mythology, the
Ganges River Dolphin is associated with Ganga, the deity of the Ganges river.
In more recent times, the
1963 Flipper movie and the subsequent popular Flipper television
series, contributed to the popularity of dolphins in Western society. The
series, created by Ivan Tors, portrayed a dolphin in a friendly relationship
with two boys, Sandy and Bud; a kind of seagoing Lassie. Flipper, a
Bottlenose Dolphin, understood English unusually well and was a marked hero. A
second Flipper movie was made in 1996, which was based on the story of
the original movie. A bottlenose dolphin also played a prominent role in the
1990's science fiction television series seaQuest DSV in which the
animal, named Darwin, could communicate with English speakers using a vocoder,
a fictional invention that translated the clicks and whistles to English and
back. More well known from this time period is probably the movie Free Willy
however, which made famous the Orca playing Willy, Keiko. The 1977 horror movie
Orca paints a less friendly picture of the animal. Here, a male Orca
takes revenge on fishermen after the killing of his mate. The 1973 movie The
Day of the Dolphin also has a dark role for dolphins, which are trained to
perform an assassination.
The renewed popularity of
dolphins in the 1960's resulted in the appearance of many dolphinariums around
the world, which have made dolphins accessible to the public. Though criticism
and more strict animal welfare laws have forced many dolphinariums to close
their doors, hundreds still exist around the world attracting large amount of
visitors. In the United States, best known are the SeaWorld marine mammal
parks, and their common Orca stage name Shamu, which they have
trademarked, is used as a synonym for Orca by some Americans. Southwest
Airlines, an American airline, has even painted three of their Boeing 737
aircraft in Shamu colors as an advertisement for the parks and have been
flying with such a livery on various aircraft since 1988.
Dolphins are also common in
contemporary literature, especially science fiction novels. For example, in the
Pern novels by Anne McCaffrey, human colonists of the fictional planet Pern
bring genetically-engineered dolphin volunteers as additional colonists. In the
second book of the Rifters trilogy, Maelstrom, by Peter Watts, enhanced
Bottlenose Dolphins are used as underwater hunter-killers by a private
military. An other military role for dolphins is found in William Gibson's
short story Johnny Mnemonic, in which cyborg dolphins are used in
war-time by the military to find submarines and, after the war, by a group of
revolutionaries to decode encrypted information. In the book Startide Rising
by author David Brin, a spaceship named Streaker is also manned by
enhanced dolphins, whose intelligence matches that of humans. More humourus is The
Hitchhiker's Guide to the Galaxy, in which dolphins are the second most
intelligent creatures on Earth, after mice, and tried in vain to warn humans of
the impending destruction of the planet. However, their behavior was
misinterpreted as playful acrobatics. Their story is told in So Long, and
Thanks for All the Fish. In the science fiction comic The Ballad of Halo
Jones, dolphins are a respected species that are the best at piloting
spacecraft, and are very peaceful and anti-war. Similarly, The Illuminatus!
trilogy features a very educated dolphin with a mature attitude as a
significant supporting character. A role for dolphins in Japanese manga is
found in Triton of the Sea (or Blue Triton) created by Osamu
Tezuka in the early 1970's. In this manga, dolphins aid the protagonists to
fight an evil would-be dictator.
Dolphins also appear
frequently in non-science fiction literature however. In the book The Music
of dolphins by author Karen Hesse, a girl is raised by dolphins from the
age of four until she is discovered by the coast guard. Fantasy author Ken
Grimwood wrote dolphins into his 1995 novel Into the Deep about a marine
biologist struggling to crack the code of dolphin intelligence, including
entire chapters written from the viewpoint of his dolphin characters. In this
book, humans and dolphins are capable of communicating via telepathy.
In the music video for Guns
N' Roses' 1993 single "Estranged", Axl Rose swims with dolphins after
plunging into the water. They end up saving his life, and he swims with the
dolphins and rides on top of one of them, and they carry him back to the top of
the water.
Occasionally, dolphins make
an appearance in computer games. Best known is the Ecco the Dolphin game
series. The games are named after their main character, Ecco, a young
Bottlenose Dolphin. The Ecco the Dolphin games hinge on the idea that cetaceans
are sapient beings and have their own underwater society. In Command and
Conquer: Red Alert 2, the Allied side can train dolphins equipped with a
sonar weapon.
An American National
Football League (NFL) team is named the Miami Dolphins. Their logo depicts an
aqua-colored Bottlenose Dolphin wearing an American football helmet and jumping
in front of a coral-colored sunburst.
http://en.wikipedia.org/w/index.php?title=Dolphin&action=history
http://www.gnu.org/copyleft/fdl.html